The overarching objective of this project is to determine the micro-scale mechanisms driving hot-spot and hot-moment carbon dynamics, for improving predictions of macro-scale carbon balance. This multi-scale interdisciplinary project will transform our knowledge of fundamental plant-soil-microbial interactions that govern past and projected carbon cycle dynamics in permafrost ecosystems, while advancing knowledge of the key biogeochemical consequences of permafrost thaw over space (i.e., plot to landscape) and time (i.e., seasonal to decadal).